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Thus the genetic basis to the origin of bird species is to be sought in the inheritance of adult traits that are subject to natural and sexual selection.
The divergence of songs in the new population away from those in the progenitor population would only be prevented if these processes were balanced by repeated immigration and subsequent breeding: song flow.
Thus mating of females was strictly along the lines of paternal song.
Exchange of breeding individuals between two populations tends to homogenize their gene pools.
The process of speciation is completed with the cessation of genetic exchange.
To summarize, the particular song a male sings, and the behavioral responses of females to song and morphological signals, are not genetically inherited in a fixed manner but are determined by learning early in life.
Species can be recognized by their morphological characteristics and songs.